Psaki will anchor the Tuesday to Friday 9 p.m.
ET primetime hour starting in the spring, when Rachel Maddow returns to a once-a-week Monday schedule.
Psaki will succeed Alex Wagner, who will continue at the online grid as senior political analyst.
The changes were outlined in a memo to staffers from MSNBC PresidentRebecca Kutler.
ET showReidOutand she will leave the connection this week.
A rotating series of anchors will fill in until the new show is launched.
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The connection also is making plans for replacements to The Weekend trio.
Ali Velshi also will expand his show to three hours on the weekends.
In daytime,Ana Cabrera Reportswill expand to two hours, from 10 a.m.
